# Tortoise Bug (*Eurygaster testudinaria*)

The Tortoise Bug is a small, round insect that looks like a little tortoise! A big shield covers its back. It sips sap from grasses, sedges and rushes.

- **Category:** [Insects](https://www.snappit.app/explore/categories/insect)
- **Rarity:** Common
- **Danger:** 1/5 · Very low
- **Habitat:** Damp and dry grassland

![Tortoise Bug](https://commons.wikimedia.org/w/index.php?title=Special:Redirect/file/Eurygaster_testudinaria_-_20070807.jpg&width=800)

Photo: Kulac at de.wikipedia., CC BY-SA 2.0 de · [Wikimedia Commons](https://commons.wikimedia.org/wiki/File:Eurygaster_testudinaria_-_20070807.jpg)

## Appearance

The Tortoise Bug has a unique, highly convex, shield-shaped body that strikingly resembles a tiny tortoise shell. Its mottled brown, grey, or yellowish colors look a lot like soil or dry vegetation. A large scutellum covers most of its back, giving it a smooth, armored look.

## Taxonomy

- **Kingdom:** Animalia
- **Phylum:** Arthropoda
- **Class:** Insecta
- **Order:** Hemiptera
- **Family:** Scutelleridae
- **Genus:** Eurygaster

## Tortoise bug, up close

The tortoise bug (Eurygaster testudinaria) is a 9–11 mm shield-backed bug (family Scutelleridae) found from Portugal and Britain across Europe and Asia to Japan. A huge plate called the scutellum covers almost its whole back, like a tortoise shell. It sucks sap from grasses, sedges and rushes in damp and dry grassland, and spends the winter as an adult in dry leaf litter.

Also called: Tortoise shieldbug, Schildkrötenwanze (German: “tortoise bug”), Gewone pantserwants (Dutch: “common armour bug”), Starrbärfis (Swedish: “sedge shieldbug”).

A shield over its back, a beak for sipping grass sap and a smelly defence. Spin it, tap it, then test yourself.

### Special abilities

- **Tortoise Shell** (Whole back under one shield; still debated): A giant scutellum covers its abdomen and folded wings in one smooth, rounded shield. [1][2][3][4][5][13][15]
  - Shield-backed bugs are named after this greatly enlarged scutellum. Their colourful tropical cousins are called jewel bugs.
  - In the green shieldbug, from a different family, the dark tips of the wings show behind the scutellum.
  - How much the shield protects it from birds or spiders has not been tested, so treat “armour” as an idea, not a measured fact.
  - Body length: Tortoise bug 9–11 mm; Scarce tortoise bug 8.5–10 mm; Sunn pest ≈12 mm; Green shieldbug 12–13.5 mm
- **Grass Sipper** (3 plant families on the menu): It pushes its beak into grass, sedge and rush stems and sucks up the sap. [6][7][16][1]
  - Main foods: grasses (Poaceae), sedges and their kin (Cyperaceae, such as cotton-grasses, Carex and clubrushes) and rushes (Juncaceae).
  - Older nymphs and adults sometimes feed on plants of the daisy and carrot families too.
  - All shield-backed bugs eat plants; none hunt other animals.
- **Stink Shield** (2 sets of scent glands in one life): Like other true bugs, it carries scent glands that release a strong-smelling defence. [9][10][17]
  - Nymphs have scent glands on top of the abdomen. Adults use glands in the thorax that open between the middle and hind legs.
  - In its close relative Eurygaster maura, the adult scent contains (E)-2-hexenal and (E)-2-hexenyl acetate, which scientists call a typical shield-backed bug mix. The tortoise bug’s own scent has not been analysed.
  - In an African shield-backed bug, the nymph glands switch off at the final moult and the adult glands take over.
  - Bugs in the shield bug group have had scent glands for at least 165 million years.
- **Egg Rows** (14 eggs per batch (sunn pest)): Tortoise bug relatives lay their eggs in tidy batches on plants: in the sunn pest, 2 rows of 7. [13][18][14]
  - A sunn pest female usually lays 28–42 eggs in 2–3 batches, on the undersides of leaves, on stems or on weeds. The eggs hatch in 6–28 days.
  - Detailed egg counts for the tortoise bug itself are not in the sources we could read.
  - Some tropical shield-backed bugs, like Pachycoris stallii in Mexico, guard their eggs and young nymphs.
  - Egg-parasitoid wasps attack Eurygaster egg masses. In one Iraqi study, 32–55% were parasitised in April.
- **Five Moults** (5 nymph stages (sunn pest)): Nymphs grow by moulting their skin, and only the last moult gives them working wings. [4][6][13][9][19]
  - Tortoise bug nymphs feed on grasses from May to August, and new adults appear from July.
  - In the sunn pest there are 5 nymph stages over 20–45 days. Wing buds show in stage 4 (front wings) and stage 5 (hind wings).
  - The openings of the adult scent glands form quickly, during the final moult.
  - Across shield bugs, nymph colours often differ from adult colours.
- **Winter Rest** (Adult the stage that survives winter): Adults spend the winter resting in dry leaf litter, sometimes in small groups, and wake in spring. [4][12][6][7][13][20]
  - In Britain adults are seen all year, with peaks in spring and autumn either side of winter.
  - They winter in dry ground litter, often in drier places than where they feed in summer.
  - The sunn pest becomes active when the leaf litter warms to about 12–13 °C.
  - In a field test in Iraq, 66–77% of wintering adults were still alive at the end of January.
- **Many Coats** (5 colour forms; still debated): Tortoise bugs come in pale brown, brown, grey, red or blackish, with faint to bold streaks. [7][2][4][5]
  - Brown is the commonest colour. The pale and dark streaks across the back range from almost invisible to strongly contrasting.
  - Colour cannot tell it apart from the scarce tortoise bug, which varies in the same way.
  - Its browns match dry grass and litter, but no study we found has tested whether this works as camouflage.
- **Hidden Wings** (12–14 °C when sunn pests start flying): Its wings stay folded under the shield until it is warm enough to fly. [3][2][13][4]
  - In shield-backed bugs the scutellum covers the wings, so they only show at take-off.
  - The sunn pest flies into the fields once the average daily temperature reaches 12–14 °C, and later flies back to woods to spend the winter.
  - In Britain the tortoise bug used to be uncommon but is now widespread in the south and is spreading northwards.

### Body parts

- **Shield** (Whole back covered by the scutellum) [1][2][3][4]
  - The big shield is the scutellum. In shield-backed bugs (Scutelleridae) it is greatly enlarged and gives the family its name.
  - It covers the whole abdomen and the folded wings, so from above the bug looks like a tiny tortoise.
  - Just in front of it, the back corners of the pronotum (the “shoulders”) stick out a little more than in its look-alike, the scarce tortoise bug.
- **Head** (A tiny dip at the front of the head) [4][5][6]
  - The best field clue: a slight central dip at the front of the head. The scarce tortoise bug (Eurygaster maura) has a smoothly rounded head.
  - The dip forms because the middle lobe (the tylus) sits a little lower than the cheek plates on either side.
  - Even experts sometimes need to check the genitals under a microscope to be sure.
- **Antennae** (1.5× segment 2 vs segment 3) [6][2]
  - The second antenna segment is only about 1.5 times as long as the third.
  - In the scarce tortoise bug it is about twice as long, another clue for telling the two apart.
- **Beak** (Sap plant juices only) [1][6][7][8]
  - Like all shield-backed bugs it is a plant-feeder, piercing plants and sucking their juices through a beak.
  - It sips from grasses, sedges and rushes. Older nymphs and adults also visit plants of the daisy and carrot families.
  - Its relative the sunn pest injects enzymes into wheat grains while it feeds, and these break down the gluten in flour.
- **Scent glands** (165 million yrs oldest fossil scent glands in its bug group) [9][10]
  - Adult true bugs have scent glands in the thorax. Nymphs use different glands on the top of the abdomen.
  - In its close relative Eurygaster maura, the adult gland is a reservoir with 2 side glands. It opens between the middle and hind legs onto rough “evaporation areas”.
  - Fossils show that bugs in the shield bug group already had scent glands in the Jurassic, about 165 million years ago.
- **Abdomen** (4 vs 2 spines in the male’s genitals) [6][5][11]
  - Under the shield, the male’s genitals carry 4 small inner spines; the scarce tortoise bug has 2.
  - Females have a small gap between the genital plates and the segment in front, which the look-alike lacks.
  - Their DNA barcodes are so similar that the tortoise bug and the scarce tortoise bug cannot be told apart by that method.
- **Food plants** (3 main plant families) [6][7][12][4]
  - Grasses, sedges (such as cotton-grasses, Carex sedges and clubrushes) and rushes.
  - It likes damp, open to half-shaded places such as tall sedge beds and bogs, and even salty ground.
  - In Britain it was once mostly a bug of damp grassland, but it now turns up in dry grassland too.
- **Eggs** (2 rows of 7 egg batch of the sunn pest) [13][14]
  - We found no detailed egg counts for the tortoise bug itself. Its relative the sunn pest lays neat batches of 14 eggs, 2 rows of 7, on leaves and stems.
  - Tiny wasps lay their own eggs inside shield bug eggs. In Iraq, the wasp Ooencyrtus telenomicida parasitised 32–55% of the eggs of wheat-field Eurygaster in April.

### Stats

| Stat | Value | Notes |
|---|---|---|
| Length | 9–11 mm | adults; one Polish account gives 7–11 mm [4][6][2][7][13] |
| Colour forms | 5 | pale brown, brown, grey, red or blackish; brown is commonest [7][2] |
| Main food plants | 3 families | grasses, sedges and rushes [6][7] |
| Range | Portugal → Japan | also Morocco and Tunisia; north to southern Scandinavia [7][6][16] |
| Generations | 1 a year | nymphs in summer, new adults from July, adults overwinter [4][12] |
| Nymph season | May–Aug | nymphs feed on grasses [4][6] |
| Winter stage | Adult | in dry ground litter, sometimes in small groups [12][6] |
| Nymph stages | 5 | measured in the sunn pest: 20–45 days in all [13] |
| Eggs per batch | 14 | 2 rows of 7, measured in the sunn pest [13] |
| Tortoise bugs in Britain | 2 | plus a third species, regionally extinct since 1885 [12] |

### Photos

- [A brown tortoise bug, the commonest colour](https://commons.wikimedia.org/wiki/File:Eurygaster_testudinaria_-_20070807.jpg) (photo: Kulac (de.wikipedia), CC BY-SA 2.0 de)
- [From above: the scutellum covers almost the whole back. Namur, Belgium](https://commons.wikimedia.org/wiki/File:Eurygaster_testudinaria_Namur%2CBelgium.jpg) (photo: Gilles San Martin, CC BY-SA 4.0)
- [A reddish-brown tortoise bug](https://commons.wikimedia.org/wiki/File:Eurygaster_testudinaria_134460476.jpg) (photo: Gilles San Martin, CC BY-SA 4.0)
- [In a bog in Dzūkija National Park, Lithuania: damp ground suits it](https://commons.wikimedia.org/wiki/File:Eurygaster_testudinaria_LT.JPG) (photo: Darius Baužys, CC BY-SA 3.0)
- [A mid-stage nymph, Worcestershire, England](https://commons.wikimedia.org/wiki/File:Tortoise_Bug._Mid_-instar_Nymph_of_Eurygaster_testudinaria_Scutelleridae_-_Flickr_-_gailhampshire_(1).jpg) (photo: gailhampshire, CC BY 2.0)
- [A mating pair, Ermingen, Germany](https://commons.wikimedia.org/wiki/File:Eurygaster-testudinaria-schildkr%C3%B6tenwanze.jpg) (photo: Holger Krisp, CC BY 3.0)
- [An adult on the Vlasina Lake plateau, Serbia](https://commons.wikimedia.org/wiki/File:Eurygaster_testudinaria_13.jpg) (photo: Slobodan Stevčić, CC BY-SA 4.0)
- [An adult among green plants, France](https://commons.wikimedia.org/wiki/File:Eurygaster_testudinaria.JPG) (photo: Jean.claude, CC BY-SA 3.0)
- [Its relative the sunn pest, a major wheat pest, in Israel](https://commons.wikimedia.org/wiki/File:Eurygaster_integriceps_in_Israel.jpg) (photo: Chaym Turak, CC BY-SA 4.0)
- [Sunn pest from above; each paper square is 5 mm](https://commons.wikimedia.org/wiki/File:Eurygaster_integriceps_-_top_(20211123_203021_-_20211123_203027_-_focus_stack).jpg) (photo: DmitTrix, CC BY 4.0)

### Questions

- **What is a tortoise bug?** The tortoise bug (Eurygaster testudinaria) is a 9–11 mm shield-backed bug of grassland in Europe and Asia. Its huge scutellum covers its back like a tortoise shell, and it sucks sap from grasses, sedges and rushes. [4][1][6]
- **Is a tortoise bug a beetle?** No. It is a true bug, with a beak for sucking plant sap. Its shield is an enlarged scutellum, not a beetle’s wing cases, and its wings are folded underneath. [1][3]
- **Are tortoise bugs harmful to people?** Tortoise bugs feed only on plant sap. Like other true bugs they can release a defensive smell, so it is best to watch them rather than pick them up, and to wash your hands if you do touch one. [1][9]
- **What do tortoise bugs eat?** Sap from grasses, sedges (such as cotton-grasses and Carex) and rushes. Older nymphs and adults sometimes feed on plants of the daisy and carrot families too. [6][7]
- **Are tortoise bugs crop pests?** Not in Europe, where they live on wild grasses, sedges and rushes. Their relative the sunn pest is a major wheat pest. Some studies in Iraq name wheat-field bugs as tortoise bugs, but these species are often confused. [6][22][14][21]
- **How can you tell a tortoise bug from the scarce tortoise bug?** Look at the front of the head: the tortoise bug has a slight central dip, while the scarce tortoise bug’s head is smoothly rounded. The tortoise bug is also slightly larger. Colour does not help, and sometimes only a microscope can tell. [4][5][6]
- **Where do tortoise bugs go in winter?** Adults spend the winter in dry ground litter, sometimes in small groups. They become active again in spring, and new adults appear from July. [12][6][4]
- **What is the sunn pest?** The sunn pest (Eurygaster integriceps) is a 12 mm relative of the tortoise bug. It feeds on wheat and barley from North Africa to Central Asia, and the enzymes it injects into grain break down the gluten in flour. [13][8][22]

### Tortoise bug myths and facts

Tap Myth or Fact for each claim, then check the answer.

- "A tortoise bug is a kind of beetle." **Myth!** It is a true bug (Heteroptera), with a beak for sucking plant sap and scent glands. Its shield is the scutellum, not a beetle’s hard wing cases. [1][9][2]
- "The tortoise bug’s “shell” is made of its wings." **Myth!** The shell is the scutellum, a greatly enlarged plate of the body. The wings are folded underneath it. [1][3][2]
- "Tortoise bugs can make a stink." **Fact!** True bugs carry scent glands as a defence: nymphs on the abdomen, adults between the middle and hind legs. Its own scent has not been analysed, but its close relative’s has. [9][10]
- "Tortoise bugs spend the winter as adults." **Fact!** Adults overwinter in dry ground litter, sometimes in small groups, and are active again in spring. [12][6]
- "You can tell a tortoise bug from its look-alike by its colour." **Myth!** Both species vary in colour. Use the slight dip at the front of the head, the antenna segments or, for certainty, the genitals under a microscope. [4][5][6]
- "Tortoise bugs only live in damp places." **Myth!** They like wet ground such as bogs and sedge beds, but also live in meadows, fallow fields and roadsides. In Britain they now turn up in dry grassland too. [7][12][4]
- "Tortoise bugs are serious wheat pests." **Not proven!** In Europe it feeds mainly on wild grasses, sedges and rushes. The famous wheat pest is its relative the sunn pest. A few studies in Iraq name wheat-field bugs as tortoise bugs, but experts warn these look-alikes are often misidentified. [6][7][23][14][22]
- "A small bug can spoil bread." **Fact!** The sunn pest, a tortoise bug relative, injects enzymes into wheat grains. They break down the gluten, so flour from damaged grain makes poor dough. [8][24][21]

### Tortoise bug records

Superlatives for the tortoise bug and its relatives, each checked against its source.

- Biggest back-plate: Shield-backed bugs (Scutelleridae), Whole abdomen (their greatly enlarged scutellum covers the abdomen and wings) [1][2]
- Hardest twins to tell apart: Tortoise bug and scarce tortoise bug, Same barcode (almost indiscernible by shape, and their DNA barcodes do not separate them) [11]
- Worst wheat bug of its region: Sunn pest (Eurygaster integriceps), 50–90% (estimated wheat yield losses in West and Central Asia and Eastern Europe) [21][22]
- Costliest bug control: Sunn pest, $42 million+ (spent on its control in West Asia) [21]
- Oldest bug stink: Scent glands of shield bugs and their kin (Pentatomomorpha), 165 million years (earliest fossil evidence, from the Jurassic) [9]

### A year in the life of a tortoise bug

One generation a year: adults sleep through winter, nymphs grow up in summer.

- **Winter**: Resting in the litter. Adults rest in dry ground litter, sometimes in small groups, often in drier spots than their summer home. [6][7][12]
- **Early spring**: Warming up. Adults become active again as the ground warms. In the sunn pest, this starts when leaf litter reaches about 12–13 °C. [12][13]
- **Spring**: First adult peak. Adults are most often seen in spring. In the sunn pest, spring is the time for breeding. [12][25]
- **Late spring**: Eggs on the plants. Relatives such as the sunn pest lay batches of 14 eggs, 2 rows of 7, on leaves and stems. They hatch in 6–28 days. [13]
- **May–August**: Nymphs feeding. Nymphs suck sap from grasses and moult as they grow. [4][6]
- **From July**: New adults. The last moult brings the full shield and working wings. [4][6]
- **Autumn**: Second peak, then shelter. Adults peak again in autumn before settling into their winter shelters. [12]

### Tortoise bug vs green shieldbug vs sunn pest

Two tortoise-shaped Eurygaster cousins and a familiar garden shieldbug from another family.

| | Tortoise bug | Green shieldbug | Sunn pest |
|---|---|---|---|
| Scientific name [26][15][13] | Eurygaster testudinaria | Palomena prasina | Eurygaster integriceps |
| Family [26][15][25] | Scutelleridae (shield-backed bugs) | Pentatomidae | Scutelleridae (shield-backed bugs) |
| Length [4][15][13] | 9–11 mm | 12–13.5 mm | ≈12 mm |
| Colour [7][15][13] | Brown, grey, reddish, pale or blackish | Bright green, bronze-brown before winter | Variable, usually light brown |
| Food [6][15][13] | Grasses, sedges, rushes | Many deciduous trees and shrubs (nymphs) | Wheat, also barley and oats |
| Winter [12][15][13] | Adult in dry litter | Adult, hibernating | Adult in leaf litter in woods |
| Generations a year [4][15][25] | 1 | 1 | 1 |
| Wings [2][15][3] | Hidden under a giant scutellum | Dark wing tips show behind the shield | Hidden under a giant scutellum |
| Wheat pest? [14][22][15][21] | Disputed: only some reports from Iraq | No: it feeds on trees and shrubs | Yes, a major one |

Tortoise bugs and sunn pests are so alike that experts sometimes need a microscope or DNA to tell Eurygaster species apart.

### Is the tortoise bug endangered?

No. It is common in much of Europe, and in Britain it is spreading.

- **IUCN Red List:** Not Evaluated (NE) (Trend unknown). It is common in Poland and widespread in central Europe. In Britain it was once uncommon, but it is now widespread in the south and expanding northwards. [27][12][7][6][4]
- Not assessed by the IUCN (Great Britain: Least Concern)
- **Threat: Natural enemies** (32–55%): Parasitic tachinid flies (Clytiomya continua and Gymnosoma nitens) attack tortoise bugs, and tiny wasps lay eggs inside Eurygaster eggs: in one Iraqi study, 32–55% of eggs were parasitised in April. [7][14]
- **Threat: Mistaken identity**: Tortoise bugs look almost the same as wheat-pest relatives. Experts warn that identification mistakes give plant protection services wrong information. [22][11]
- **Helping: Spreading north**: Once uncommon in Britain, it is now widespread in southern Britain and Ireland and is expanding its range northwards. [4]
- **Helping: Volunteer recording**: Records sent to the Terrestrial Heteroptera Recording Scheme feed the British atlas of shieldbugs. [12]
- **Helping: Smarter pest control**: For the sunn pest, scientists measure nymph numbers and egg parasitism by wasps to target control where it is really needed. [28]
- **Scarce tortoise bug (Eurygaster maura)** (NE): Nationally Scarce in Britain, mostly on chalk and other lime-rich grassland in southern England. [12][5]
- **Eurygaster austriaca** (NE): Regionally extinct in Britain: it lived on the Kent coast but has not been recorded there since 1885. [12]
- **Sunn pest (Eurygaster integriceps)** (NE): Far from rare: it is the most important wheat pest in Russia and a major pest from North Africa to the Middle East and Eastern Europe. [22][8]

### Looks

- **Brown**: The commonest colour: brown with paler and darker streaks that can be faint or bold. (Most tortoise bugs you will meet) [7][2]
- **Reddish brown**: Some tortoise bugs are red-brown or reddish. Others are grey, pale brown or almost black. (One of several colour forms) [7][2][4]
- **Nymph**: Young tortoise bugs have no working wings yet. They feed on grasses from May to August and become adults from July. (Wing buds appear in the last nymph stages (sunn pest)) [4][13][19]
- **Sunn pest (relative)**: Eurygaster integriceps: about 12 mm, usually light brown, with the same giant shield. (A major pest of wheat and barley) [13][3][21][22]

### Tortoise bug quiz

1. What is the tortoise bug’s big shield called? **Scutellum.** It is the scutellum, greatly enlarged in shield-backed bugs. [1]
2. Which family does the tortoise bug belong to? **Scutelleridae.** Scutelleridae, the shield-backed or jewel bugs. [26][1]
3. How long is an adult tortoise bug? **9–11 mm.** Adults are 9–11 mm long. [4][6]
4. What does it mostly feed on? **Grasses, sedges and rushes.** It sucks sap from grasses, sedges and rushes. [6][7]
5. How does it spend the winter? **As an adult in dry litter.** Adults overwinter in dry ground litter, sometimes in small groups. [12][6]
6. Where do an adult shield bug’s scent glands open? **Between the middle and hind legs.** In Eurygaster maura they open between the middle and hind legs. [10]
7. What is the best clue for telling it from the scarce tortoise bug? **A slight dip at the front of the head.** The tortoise bug has a slight central dip at the front of the head; colour varies in both. [4][5]
8. Which relative is a major wheat pest? **Sunn pest.** The sunn pest, Eurygaster integriceps. [22][21]
9. How does sunn pest feeding spoil flour? **Enzymes break down the gluten.** Enzymes it injects into wheat grains break down gluten, so the dough is poor. [8][24]
10. How many eggs are in a sunn pest egg batch? **14.** 14 eggs: 2 neat rows of 7. [13]
11. When do tortoise bug nymphs feed? **May–August.** Nymphs feed on grasses from May to August. [4]
12. How old are the oldest fossil scent glands of shield bugs and their kin? **165 million years.** About 165 million years, from the Jurassic. [9]

### Sources (checked 2026-10-16)

1. [Wu YZ, Rédei D, Eger J, et al. (2018). Phylogeny and the colourful history of jewel bugs (Insecta: Hemiptera: Scutelleridae). Cladistics 34(5): 502–516.](https://doi.org/10.1111/cla.12224)
2. [Wikipedia (Dutch). Gewone pantserwants (Eurygaster testudinaria).](https://nl.wikipedia.org/wiki/Gewone_pantserwants)
3. [Wikipedia (English). Eurygaster integriceps (sunn pest).](https://en.wikipedia.org/wiki/Eurygaster_integriceps)
4. [British Bugs (T. Bantock & J. Botting). Eurygaster testudinaria (Scutelleridae), Tortoise Bug.](https://www.britishbugs.org.uk/heteroptera/Scutellerideae/e_testudinaria.html)
5. [British Bugs (T. Bantock & J. Botting). Eurygaster maura (Scutelleridae), Scarce Tortoise Bug.](https://www.britishbugs.org.uk/heteroptera/Scutellerideae/e_maura.html)
6. [Wikipedia (German). Schildkrötenwanze (Eurygaster testudinaria), summarising Wachmann, Melber & Deckert (2008), Wanzen vol. 4.](https://de.wikipedia.org/wiki/Schildkr%C3%B6tenwanze)
7. [Wikipedia (Polish). Żółwinek żółwik (Eurygaster testudinaria), summarising Lis, Lis & Ziaja (2012), Heteroptera Poloniae 2.](https://pl.wikipedia.org/wiki/%C5%BB%C3%B3%C5%82winek_%C5%BC%C3%B3%C5%82wik)
8. [Yandamuri RC, Gautam R, Darkoh C, Dareddy V, El-Bouhssini M, Clack BA (2014). Cloning, expression, sequence analysis and homology modeling of the prolyl endoprotease from Eurygaster integriceps Puton. Insects 5(4): 762–782.](https://doi.org/10.3390/insects5040762)
9. [Dai R, Du S, Ren D, Engel MS, Yao Y (2025). A foul odor from the Mesozoic: early evolution of scent gland defense in pentatomomorphan true bugs. BMC Biology 23: 345.](https://doi.org/10.1186/s12915-025-02446-x)
10. [Durak D, Kalender Y (2007). Fine structure and chemical analysis of the metathoracic scent gland of Eurygaster maura (Heteroptera: Scutelleridae). Folia Biologica (Kraków) 55(3): 133–141.](https://doi.org/10.3409/173491607781492551)
11. [Syromyatnikov MY, Golub VB, Kokina AV, Soboleva VA, Popov VN (2017). DNA barcoding and morphological analysis for rapid identification of most economically important crop-infesting Sunn pests belonging to Eurygaster. ZooKeys 706: 51–71.](https://doi.org/10.3897/zookeys.706.13888)
12. [Bantock T. (2018). Provisional atlas of shieldbugs and allies, v1.1. Terrestrial Heteroptera Recording Scheme.](https://www.britishbugs.org.uk/Provisional_atlas_of_shieldbugs_and_allies_2018.pdf)
13. [AgroAtlas (Interactive agricultural ecological atlas of Russia and neighbouring countries). Eurygaster integriceps Puton, sunn pest.](https://www.agroatlas.ru/en/content/pests/Eurygaster_integriceps/)
14. [Al-Hadrawi TI, AllAllah AJ, Abdullah LH (2023). The relationship between the presence of sunn pest Eurygaster testudinaria Geoffroy and its egg parasitoids in Najaf Province. IOP Conference Series: Earth and Environmental Science 1158: 072017.](https://doi.org/10.1088/1755-1315/1158/7/072017)
15. [British Bugs (T. Bantock & J. Botting). Palomena prasina (Pentatomidae), Green Shieldbug.](https://www.britishbugs.org.uk/heteroptera/Pentatomidae/palomena_prasina.html)
16. [Wikipedia (English). Eurygaster testudinaria.](https://en.wikipedia.org/wiki/Eurygaster_testudinaria)
17. [Hamilton JGC, Gough AJE, Staddon BW, Games DE (1985). Multichemical defense of plant bug Hotea gambiae (Heteroptera: Scutelleridae). Journal of Chemical Ecology 11(10): 1399–1409.](https://doi.org/10.1007/BF01012140)
18. [Williams L, Coscarón MC, Dellapé PM, Roane TM (2005). The shield-backed bug, Pachycoris stallii: description of immature stages, effect of maternal care on nymphs, and notes on life history. Journal of Insect Science 5: 29.](https://doi.org/10.1093/jis/5.1.29)
19. [Medina I, Vega-Trejo R, Wallenius T, et al. (2020). No link between nymph and adult coloration in shield bugs: weak selection by predators. Proceedings of the Royal Society B 287: 20201011.](https://doi.org/10.1098/rspb.2020.1011)
20. [Al-Hadrawi TI, AllAllah AJ (2023). Effect of overwintering time on survival adults of Eurygaster testudinaria Geoffroy in Iraq. IOP Conference Series: Earth and Environmental Science 1158: 072008.](https://doi.org/10.1088/1755-1315/1158/7/072008)
21. [University of Vermont Entomology Research Laboratory. Sunn Pest: unlocking the mysteries of an ancient problem.](https://www.uvm.edu/~entlab/sunnpest/)
22. [Neimorovets V. (2020). Review of the genus Eurygaster (Hemiptera: Heteroptera: Scutelleridae) of Russia. Zootaxa 4722(6).](https://doi.org/10.11646/zootaxa.4722.6.1)
23. [Salis L, Goula M, Izquierdo J, Gordún E (2013). Population density and distribution of wheat bugs infesting durum wheat in Sardinia, Italy. Journal of Insect Science 13: 50.](https://doi.org/10.1673/031.013.5001)
24. [Dolgikh V, Tsarev A, Timofeev S, et al. (2020). Antibodies raised against a Sunn bug (Eurygaster integriceps) recombinant protease, rGHP3p2, can inhibit gluten-hydrolyzing activity. Food Science & Nutrition 8(1): 703–708.](https://doi.org/10.1002/fsn3.1361)
25. [Alborzi Z, Amiri A, Piulachs MD, Bandani AR (2025). Tissue-specific transcriptomic and endocrine signatures of obligatory reproductive diapause in Eurygaster integriceps. Scientific Reports 15: 42999.](https://doi.org/10.1038/s41598-025-27154-y)
26. [GBIF Backbone Taxonomy. Eurygaster testudinaria (Geoffroy, 1785), species 5758794.](https://www.gbif.org/species/5758794)
27. [Wikidata. Eurygaster testudinaria (Q1130881).](https://www.wikidata.org/wiki/Q1130881)
28. [Duman M, Alaserhat İ (2025). Assessment of Eurygaster integriceps parameters to improve integrated pest management in Southeastern Türkiye wheat fields. Scientific Reports 15: 26031.](https://doi.org/10.1038/s41598-025-10340-3)

## Measurements & details

- **Wingspan:** 15-25 mm
- **Size:** 7-11 mm
- **Lifespan:** 180-300 days
- **Leg Count:** 6

## Diet & Feeding

These tiny bugs drink sap from grasses, sedges and rushes.

- **Primary Foods:** Grasses (Poaceae), Sedges (Cyperaceae), Rushes (Juncaceae)
- **Foraging Method:** Foraging

## Ecological connections

- **Depends on:** [Grasses](https://www.snappit.app/explore/grasses) (*Poaceae*) — Adults and nymphs feed on the sap of grasses.
- **Depends on:** Sedges (*Cyperaceae*) — They pierce sedge stems and sip the sap.
- **Parasite:** [Tachinid flies](https://www.snappit.app/explore/tachinid-flies) (*Clytiomya continua*) — These parasitic flies attack Tortoise Bugs.

## Traits

- [Solitary](https://www.snappit.app/explore/traits/solitary): Solitary animals live alone for most of their lives, only interacting with others for mating or parental care.
- [Camouflaged](https://www.snappit.app/explore/traits/camouflaged): Camouflaged describes organisms that possess coloring or patterns that allow them to blend into their environment.
- [Herbivorous](https://www.snappit.app/explore/traits/herbivorous): This trait characterizes organisms whose diet consists entirely or primarily of plant material.
- [Armored](https://www.snappit.app/explore/traits/armored): Armored describes animals protected by a hard, external covering or shell for defense.
- [Diurnal](https://www.snappit.app/explore/traits/diurnal): Diurnal animals are primarily active during daylight hours, typically resting or sleeping at night.

## Collections

- [Insects](https://www.snappit.app/explore/collections/insects)

## Safety

**Danger:** 1/5 · Very low

It's best to observe Tortoise Bugs from a distance and not touch them.

## Frequently Asked Questions

### How to identify Tortoise Bug?

Look for six legs, antennae, three body sections and the shape of the wings. Beetles have hardened front wings; flies have just one working pair. Tiny insects may need a close view before their group is clear. Insect Identifier: upload a photo and compare the visible details.

### What is Tortoise Bug's wingspan?

15-25 mm

### How big is Tortoise Bug?

7-11 mm

### How long does Tortoise Bug live?

180-300 days

### What does Tortoise Bug eat?

These tiny bugs drink sap from grasses, sedges and rushes.

### Where is Tortoise Bug usually found?

Damp and dry grassland

### How many legs does Tortoise Bug have?

6

## More Insects

- [Common Eastern Bumble Bee](https://www.snappit.app/explore/common-eastern-bumble-bee) (*Bombus impatiens*)
- [Spotted Lanternfly](https://www.snappit.app/explore/spotted-lanternfly) (*Lycorma delicatula*)
- [Brown-belted Bumble Bee](https://www.snappit.app/explore/brown-belted-bumble-bee) (*Bombus griseocollis*)
- [Eastern Carpenter Bee](https://www.snappit.app/explore/eastern-carpenter-bee) (*Xylocopa virginica*)

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Source: [Tortoise Bug: Facts & Places to See — Snappit](https://www.snappit.app/explore/tortoise-bug)