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Glowworm beetle

Phengodidae

Glowworm beetles (family Phengodidae), also called railroad worms, are soft-bodied beetles found almost only in the Americas. Their larvae and wingless, larva-shaped adult females glow from pairs of lights along the body, while the small winged males have feathery antennae. Railroad worms of the genus Phrixothrix also carry a red light on the head: the only true red light known from a land animal.

Also known as Railroad worm, Phengodid beetle

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Glowworm beetle
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Glowworm beetle

Rarity: Common

Female (railroad worm)

Glowworm beetle abilities

  • 11 pairs steady side lights

  • 628 nm reddest beetle light

  • 2 hollow, sickle-shaped jaws

  • 24 lights on a PhrixothrixStill debated

  • 12 segments in each male antenna

  • 3 vs 12 antenna segments, female vs male

  • 15 in 100 reactions give off red light

  • 53 eggs laid by one female

Real glowworm beetles

Photos from Wikimedia Commons. Some of them can repaint the 3D model.

  • A railroad worm shown with its lights switched on and with its lights off
    Female (railroad worm)Larva-shaped for life: a long, soft, segmented body with green-yellow lights along the sides and, in Phrixothrix, red lights on the head.Wingless adult · glows · lays eggsLights on, lights off: the two looks of a railroad worm. Photo: Aaron Pomerantz, CC BY 2.0
  • A Phrixothrix hirtus railroad worm in the Brazilian Atlantic Forest
    Female (railroad worm)Larva-shaped for life: a long, soft, segmented body with green-yellow lights along the sides and, in Phrixothrix, red lights on the head.Wingless adult · glows · lays eggsPhrixothrix hirtus, the red-light railroad worm, in Brazil. Photo: Thomas Fuhrmann, CC BY-SA 4.0
  • A larva-shaped Phengodes glowworm beetle, black with rows of orange spots, on sandy ground
    LarvaAlmost identical to the female, but with no egg opening. It grows to 15–65 mm before turning into a pupa.Hunts millipedes · glowsA Phengodes in Missouri: a female or a larva. Photo: Andy Reago & Chrissy McClarren, CC BY 2.0
  • A larva-shaped Phengodes glowworm beetle
    Female (railroad worm)Larva-shaped for life: a long, soft, segmented body with green-yellow lights along the sides and, in Phrixothrix, red lights on the head.Wingless adult · glows · lays eggsFemale or larva? Experts check for the tiny egg opening. Photo: Benjamin Smith, CC BY 2.0
  • A western banded glowworm, a larva-shaped beetle with dark bands
    Western banded glowworm (Zarhipis integripennis)Photo: nancybeetoo, CC BY 2.0
  • A male Phengodes glowworm beetle with feathery antennae and short wing covers
    Winged maleA small brown-to-black beetle, about 6–35 mm long, with short wing covers and big feathery antennae.Flies at night · follows scent · short-livedA winged male: note the feathery antennae. Photo: Insects Unlocked, CC0
  • A wingless female European glow-worm
    European glow-wormA different beetle: the wingless female of a firefly, Lampyris noctiluca. She glows steadily at night to attract flying males.Firefly family (Lampyridae) · larvae eat snailsThe European glow-worm is a firefly, not a railroad worm. Photo: Tero Keski-Valkama, CC0
  • A European glow-worm (Lampyris noctiluca)
    European glow-wormA different beetle: the wingless female of a firefly, Lampyris noctiluca. She glows steadily at night to attract flying males.Firefly family (Lampyridae) · larvae eat snailsEuropean glow-worm in the UK. Photo: AJC1, CC BY-SA 2.0

Glowworm beetle by the numbers

Measured by scientists in the field and in the lab.

Species
≈250
244 counted in 2023, almost all in the Americas⁠[2, 7]
Side lights
11 pairs
in Phrixothrix, plus 2 red head lights⁠[1]
Head light colour
628 nm
deep red: the reddest beetle light measured⁠[1]
Side light colour
535–565 nm
green to yellow in Phrixothrix⁠[3]
Larva length
15–65 mm
full-grown larva, depending on species⁠[4]
Male length
6–35 mm
small winged beetles⁠[4]
Eggs
53
laid by one Phengodes female (1906 record)⁠[4]
Time as a pupa
12–35 days
western banded glowworm: 12–13 days for females, 20–35 for males⁠[4]
Genome size
≈3.4 Gb
Phrixothrix hirtus: the largest known in its beetle superfamily⁠[2]
Red light efficiency
15%
of reactions give off light, vs over 40% for green beetle enzymes⁠[14]

Glowworm beetle myths and facts

The word glowworm causes a lot of mix-ups. Test what you know.

0 of 8 answered

  1. “Glowworms are a kind of worm.”

  2. “Every animal called a glowworm is the same kind of creature.”

  3. “Railroad worms flash on and off like fireflies.”

  4. “The railroad worm's red head light is the only red light made by a land animal.”

  5. “Male glowworm beetles find females by following their glow.”

  6. “Glowworm beetle larvae eat snails.”

  7. “Male glowworm beetles never glow.”

  8. “A female railroad worm is just a baby that never grew up.”

Glowworm beetle records

Small beetles, big superlatives. Each one is backed by a study.

  • Only red light on land

    622–628 nm

    Phrixothrix hirtus

    The only land animal known to make true red bioluminescence.⁠[2, 1]

  • Widest range of light colours

    Green to red

    Railroad worms (Phengodidae)

    Their enzymes make the widest range of colours of any beetle group, from about 542 nm green to 628 nm red.⁠[8, 24, 1]

  • Biggest genome in its superfamily

    ≈3.4 Gb

    Phrixothrix hirtus

    The largest genome described among the click beetles, fireflies and their relatives (Elateroidea).⁠[2]

  • Oldest glowing relative

    ≈99 million years

    Cretophengodes azari

    A male beetle in amber from Myanmar, with a light organ on its abdomen, linking glowworm beetles to harder-bodied relatives.⁠[11]

  • First orange railroad-worm enzyme

    600 nm

    Euryopa clarindae

    In 2024 scientists described the first orange-glowing railroad worm luciferase, from males of a rare species of the Brazilian Cerrado.⁠[8]

The railroad worm's red light

Why scientists study one small Brazilian beetle's head lamp.

  • 628 nm

    reddest beetle light

    Measured from the head lights of Phrixothrix hirtus. The side lights of a related species glow green at 542 nm.⁠[1]

  • 71%

    same enzyme recipe

    The red and green luciferases share 71% of their building blocks. Small changes shift the colour.⁠[1]

  • 15 in 100

    reactions make red light

    The red enzyme is dimmer than green ones, which manage more than 40 in 100.⁠[14]

  • 9.8×

    brighter in the lab

    By changing a few building blocks, scientists made red enzymes up to 9.8 times brighter in cell extracts.⁠[23]

  • Red

    passes through tissue

    Blood and skin absorb red light less than green, so researchers use red-glowing enzymes to follow infections and disease inside living animals.⁠[15]

Railroad worm red light comes from the same luciferin fuel as a firefly's yellow-green; only the enzyme differs.

Life of a glowworm beetle

From a glowing egg to a winged male or a larva-shaped mother. Timings come from the few species that have been studied.

  1. Egg

    Laid in a group

    A mother lays her eggs together on the ground, 53 in one record, and curls around them.⁠[4]

  2. Up to 1 month

    Eggs light up

    The eggs glow, but it can take up to a month after laying before they become luminous.⁠[4]

  3. Larva

    Millipede hunter

    The larva hunts millipedes in soil and leaf litter and grows to 15–65 mm.⁠[4, 5]

  4. Pupa

    The big change

    In the western banded glowworm, a female spends 12–13 days as a pupa and a male 20–35 days.⁠[4]

  5. Adult female

    Larva shape for life

    She stays wingless and larva-like, glows, and gives off a scent that draws in males.⁠[4]

  6. Adult male

    Short night flights

    The winged male follows her scent through the dark. Males are short-lived and probably do not feed.⁠[4, 7]

  7. Mother

    Glowing guard

    After mating she curls around her new eggs, often glowing continuously, perhaps until they hatch.⁠[4]

Railroad worm vs European glow-worm vs New Zealand glowworm

Three animals, one confusing name. Only the first is a glowworm beetle.

Railroad worm (Phengodidae) vs European glow-worm (Lampyris noctiluca) vs New Zealand glowworm (Arachnocampa luminosa)
Railroad wormEuropean glow-wormNew Zealand glowworm
What it is⁠[4, 18, 21, 22]A beetle (family Phengodidae)A firefly beetle (family Lampyridae)A fly: the larva of a fungus gnat (Keroplatidae)
Who glows⁠[4, 20, 17, 22]Larvae, wingless females, and the males of some groupsAbove all the wingless adult femaleThe larva
Light colour⁠[3, 1, 18, 25, 9]Green-yellow sides (535–565 nm), red head (600–628 nm) in PhrixothrixGreen, a little bluer than an American firefly'sBlue-green (about 488 nm in an Australian relative)
Why it glows⁠[4, 17, 22]Thought to warn predators it tastes badTo attract flying malesTo lure flying insects to its sticky threads
Larva's food⁠[4, 5, 19, 26, 22]MillipedesSnailsSmall flying insects caught on sticky silk lines
Where⁠[7, 20, 17, 25]The Americas, from southern Canada to Chile (a few in Turkey and the Middle East)EuropeNew Zealand
Light chemistry⁠[3, 18, 25]Firefly-type luciferinFirefly-type luciferinA completely different luciferin

Glowworm beetles and European glow-worms are both beetles, yet genome studies suggest beetles evolved light more than once.

Are glowworm beetles endangered?

Nobody knows yet. Glowworm beetles are rarely seen, many species have not even been named, and none has been assessed for the IUCN Red List.

IUCN Red List

Not Evaluated (NE)

Trend unknown

No glowworm beetle species has an IUCN assessment, so how they are doing is unknown.⁠[27, 4]

Not on the IUCN Red List (checked October 2026)

Threats

  • Forest loss>80%

    Many Brazilian railroad worms live in the Atlantic Forest and Cerrado. More than 80% of the Atlantic Forest's remaining patches are smaller than 50 ha.⁠[20, 28]

  • Light pollution15 nights

    Artificial light at night harms many insects. In the European glow-worm, females near lights glowed for up to 15 nights without finding a mate.⁠[29, 30]

  • Too little known≈100

    Around 100 species may be waiting in museum collections to be named. It is hard to protect animals nobody has described.⁠[4]

What’s helping

  • Darker nights

    Switching off or dimming outdoor lights helps night insects. In glow-worm experiments, red and yellow lamps disturbed mating less than white or blue ones.⁠[29, 31]

  • Naming and studying

    Scientists are still describing species and using DNA to match larvae with their adults, which had been done for only one Brazilian species before 2025.⁠[20]

  • Look, don't take

    Females cannot fly and move only short distances, so local groups depend on the females staying where they are.⁠[6]

Glowing relatives

  • European glow-worm (Lampyris noctiluca)Near Threatened

    Listed as Near Threatened. Artificial light makes it harder for females to attract mates.⁠[27, 17, 30]

  • North American firefliesVaries by species

    Of 132 species assessed, at least 18 (14%) are threatened, mainly by habitat loss, light pollution and climate change.⁠[32]

Glowworm beetle quiz

12 questions. Every answer is somewhere on this page.

Question 1 of 12Score 0

What kind of animal is a glowworm beetle?

Glowworm beetle field notes

Diet, food web, safety and badges from the Snappit field guide.

Diet & Feeding

Glowworm larvae are tiny hunters. As far as scientists know, they hunt millipedes.

Primary Foods

  • Millipedes

Age differences: Adult males generally do not eat, while larvae are active predators.

Ecological connections

  • American MillipedeHunts

    Glowworm larvae prey on these millipedes on the forest floor.

  • Masked ShrewPossible predator (unconfirmed)

    Shrews might eat glowworm larvae that live in leaf litter, but this has not been confirmed.

  • American RobinPossible predator (unconfirmed)

    Ground-feeding birds like robins might eat glowworm larvae, but this has not been confirmed.

Safety

Danger 1/5 · Very low

Avoid touching or disturbing glowworms in their natural habitat to ensure they can continue their important role in the ecosystem.

Traits

Collections

Classification

Kingdom
Animalia
Phylum
Arthropoda
Class
Insecta
Order
Coleoptera
Family
Phengodidae

Sources for these glowworm beetle facts

32 studies and references · checked
  1. Viviani VR, Bechara EJH, Ohmiya Y (1999). Cloning, sequence analysis, and expression of active Phrixothrix railroad-worms luciferases: relationship between bioluminescence spectra and primary structures. Biochemistry 38:8271–8279.
  2. Amaral DT, Mitani Y, Bonatelli IAS, Cerri R, Ohmiya Y, Viviani VR (2023). Genome analysis of Phrixothrix hirtus (Phengodidae) railroad worm shows the expansion of odorant-binding gene families and positive selection on morphogenesis and sex determination genes. Gene 850:146917.
  3. Viviani VR, Bechara EJH (1993). Biophysical and biochemical aspects of phengodid (railroad-worm) bioluminescence. Photochemistry and Photobiology 58:615–622.
  4. Branham M (2005, revised 2019). Glow-worms, railroad-worms (Insecta: Coleoptera: Phengodidae). UF/IFAS Featured Creatures EENY332 / EDIS IN609.
  5. Eisner T, Eisner M, Attygalle AB, Deyrup M, Meinwald J (1998). Rendering the inedible edible: circumvention of a millipede's chemical defense by a predaceous beetle larva. PNAS 95:1108–1113.
  6. Richmond M, Caterino MS (2012). Population subdivision of Zarhipis integripennis (Coleoptera: Phengodidae) in southern California. Annals of the Entomological Society of America 105:241–252.
  7. Wikipedia contributors. Phengodidae. English Wikipedia (accessed 2026-10-16).
  8. Viviani VR, Benites GR, Souza DR, Pelentir GF, Reis RM, Bechara EJH, Tomazini A (2024). The orange light emitting luciferase from the rare Euryopa clarindae adult railroadworm (Coleoptera: Phengodidae). Photochemical & Photobiological Sciences 23.
  9. Lee J (1976). Bioluminescence of the Australian glow-worm, Arachnocampa richardsae Harrison. Photochemistry and Photobiology 24:279–285.
  10. Wikipedia contributors. Railroad worm. English Wikipedia (accessed 2026-10-16).
  11. Li YD, Kundrata R, Tihelka E, Liu Z, Huang D, Cai C (2021). Cretophengodidae, a new Cretaceous beetle family, sheds light on the evolution of bioluminescence. Proceedings of the Royal Society B 288:20202730.
  12. He J, Li J, Zhang R, et al. (2024). Multiple origins of bioluminescence in beetles and evolution of luciferase function. Molecular Biology and Evolution 41:msad287.
  13. Fallon TR, Lower SE, Chang CH, et al. (2018). Firefly genomes illuminate parallel origins of bioluminescence in beetles. eLife 7:e36495.
  14. Bevilaqua VR, Matsuhashi T, Oliveira G, Oliveira PSL, Hirano T, Viviani VR (2019). Phrixotrix luciferase and 6'-aminoluciferins reveal a larger luciferin phenolate binding site and provide novel far-red combinations for bioimaging purposes. Scientific Reports 9:8998.
  15. Viviani VR, Bevilaqua VR, de Souza DR, Pelentir GF, Kakiuchi M, Hirano T (2020). A very bright far-red bioluminescence emitting combination based on engineered railroad worm luciferase and 6'-amino-analogs for bioimaging purposes. International Journal of Molecular Sciences 22:303.
  16. Viviani VR (2002). The origin, diversity, and structure function relationships of insect luciferases. Cellular and Molecular Life Sciences 59:1833–1850.
  17. Elgert C, Hopkins J, Kaitala A, Candolin U (2020). Reproduction under light pollution: maladaptive response to spatial variation in artificial light in a glow-worm. Proceedings of the Royal Society B 287:20200806.
  18. Sala-Newby GB, Thomson CM, Campbell AK (1996). Sequence and biochemical similarities between the luciferases of the glow-worm Lampyris noctiluca and the firefly Photinus pyralis. Biochemical Journal 313:761–767.
  19. Krämer J, Hölker P, Predel R (2024). How to overcome a snail? Identification of putative neurotoxins of snail-feeding firefly larvae (Coleoptera: Lampyridae, Lampyris noctiluca). Toxins 16:272.
  20. Roza AS, Souza DR, Pelentir GF, Barbosa FF, Mermudes JRM, Viviani VR (2025). Bioluminescence, morphology, biological aspects and molecular association of larvae and adults of Stenophrixothrix (Coleoptera: Phengodidae). Zootaxa 5689(3).
  21. von Byern J, Chandler P, Merritt D, et al. (2019). Biomechanical properties of fishing lines of the glowworm Arachnocampa luminosa (Diptera; Keroplatidae). Scientific Reports 9:3082.
  22. Merritt DJ, Patterson R (2018). Environmental influences on the bioluminescence display of the glow-worm, Arachnocampa flava. Austral Entomology 57:107–117.
  23. Li X, Nakajima Y, Niwa K, Viviani VR, Ohmiya Y (2010). Enhanced red-emitting railroad worm luciferase for bioassays and bioimaging. Protein Science 19:26–33.
  24. Arnoldi FG, da Silva Neto AJ, Viviani VR (2010). Molecular insights on the evolution of the lateral and head lantern luciferases and bioluminescence colors in Mastinocerini railroad-worms (Coleoptera: Phengodidae). Photochemical & Photobiological Sciences 9:87–92.
  25. Watkins OC, Sharpe ML, Perry NB, Krause KL (2018). New Zealand glowworm (Arachnocampa luminosa) bioluminescence is produced by a firefly-like luciferase but an entirely new luciferin. Scientific Reports 8:3278.
  26. von Byern J, Dorrer V, Merritt DJ, Chandler P, Stringer I, Marchetti-Deschmann M (2016). Characterization of the fishing lines in Titiwai (=Arachnocampa luminosa Skuse, 1890) from New Zealand and Australia. PLoS ONE 11:e0162687.
  27. IUCN Red List checklist dataset on GBIF (searched for Phengodidae, Phrixothrix and Lampyris noctiluca, 2026-10-16).
  28. Ribeiro MC, Metzger JP, Martensen AC, Ponzoni FJ, Hirota MM (2009). The Brazilian Atlantic Forest: how much is left, and how is the remaining forest distributed? Biological Conservation 142:1141–1153.
  29. Owens ACS, Cochard P, Durrant J, Farnworth B, Perkin EK, Seymoure B (2020). Light pollution is a driver of insect declines. Biological Conservation 241:108259.
  30. Van den Broeck M, De Cock R, Van Dongen S, Matthysen E (2021). Blinded by the light: artificial light lowers mate attraction success in female glow-worms (Lampyris noctiluca L.). Insects 12:734.
  31. Kivelä L, Elgert C, Lehtonen TK, Candolin U (2023). The color of artificial light affects mate attraction in the common glow-worm. Science of the Total Environment 857:159451.
  32. Fallon CE, Walker AC, Lewis S, et al. (2021). Evaluating firefly extinction risk: initial red list assessments for North America. PLoS ONE 16:e0259379.

Frequently Asked Questions

How to identify Glowworm beetle?

Look for hard wing covers meeting in a line down the back, then compare the antennae, body shape, legs and markings. Metallic colors shift a lot with lighting. Beetle Identifier: upload a photo and compare the visible details.

How big is Glowworm beetle?

6-65 mm

How long does Glowworm beetle live?

200-700 days

What does Glowworm beetle eat?

Glowworm larvae are tiny hunters. As far as scientists know, they hunt millipedes.

Where is Glowworm beetle usually found?

Forests and dry areas

How many legs does Glowworm beetle have?

6

What is the difference between a glowworm and a firefly?

Glowworm is a name, not one animal. Glowworm beetles of the Americas (Phengodidae) are their own beetle family. The European glow-worm is actually a firefly whose females have no wings. The New Zealand glowworm is a fly larva.

Why is it called a railroad worm?

Its rows of glowing spots along the body look like the lit windows of a train at night.

Why do glowworm beetles glow?

Scientists think the glow warns night-time predators that they taste bad. Males find females by scent, not light, so the glow is probably not a love signal.

What do glowworm beetles eat?

The larvae and larva-shaped females hunt millipedes, paralysing them quickly with digestive juice from their hollow jaws. Adult males are short-lived and probably do not feed.

Where do glowworm beetles live?

Almost all live in the Americas, from southern Canada to Chile, in places from dry parts of California to tropical forests. A small group lives in Turkey and the Middle East.

Do male glowworm beetles glow?

Usually not, but males in the group that includes Phrixothrix glow all their adult life, and Stenophrixothrix males give off yellow light.

Are glowworm beetles dangerous?

There is no sign that they are a danger to people: their jaws are made for millipedes. Still, don't pick them up. When handled, a railroad worm larva may release a reddish liquid that may taste bad or irritate. Watch without touching, and wash your hands if you do touch one.

How can I see glowworm beetles in the wild?

Look on warm, damp nights on soil and leaf litter. Use a dim red light, which disturbed glow-worm mating less than white or blue light in experiments. Watch without touching and leave them where they are.

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